1. Primary scientific publications in refereed international journals

108 - Amit T, Moskovich R, Jacobi Y, Shumway SE, Ward JE, Beninger PG, Yahel G,Loya Y. 2023. Feeding on the smallest cells: an in situ study of picoplankton capture by bivalve molluscs from oligotrophic waters
Front. Mar. Sci. 10:1184773  pdf

107 - Beninger PG, Chérel D, Le Pennec G, Shumway SE. 2022. Evolutionary and ecological insights from vital staining of bivalve oocytes: A Red Queen at the sweepstakes?
Ecosphere. 62/166   pdf

106 - Eveillard-Buchoix M, Beninger PG. 2022. Between a rock and a large place: the importance of multi-scale geomorphological features to seabird nest site selection.
Ecology. 103(1)   pdf

105 - Beninger PG , Chérel D, Kessler L. 2021. Examining bivalve fecundity: oocyte viability revealed by Neutral Red vital staining.
Aquacult Int.   pdf

104 - Chérel D, Beninger PG , Le Pennec G. 2020. Two enigmas may solve each other: the oocyte coat and atresia in the common cockle, Cerastoderma edule (Linnaeus, 1758).
Marine Biology. 167:104   pdf

103 - Beninger PG , ROBERT W. ELNER. 2020. On the tip of the tongue: natural history observations that transformed shorebird ecology
Ecosphere. 11(5):1-4   pdf

102 - Chérel D, Beninger PG . 2019. Oocyte atresia and its effect on reproductive effort of the Common Cockle Cerastoderma edule (Linneaus, 1758).
J. Shellfish Res. 38: 603-609   pdf

101 - Beninger PG , Chérel D. 2019. Cloaked bivalve oocytes: lessons in evolution, ecology, and scientific awareness.
Ecology 100 (12)  pdf

100 - Eveillard-Buchoux M, Beninger PG , Chadenas C, Sellier D. 2019. Small-scale natural landscape features and seabird nesting sites: the importance of geodiversity for conservation.
Landscape Ecol 34: 2295–2306   pdf

99 - Beninger PG , Backeljau T. 2019. Understanding taxonomic and nomenclatural instability – a case study of the Manila clam.
Aquaculture 504: 375-379   pdf

98 - Chérel D., Beninger PG . 2017. Oocyte Atresia Characteristics and Effect on Reproductive Effort of Manila Clam Tapes philippinarum (Adams and Reeve, 1850).
Journal of Shellfish Research 36(3): 549-557   pdf

97 - Eveillard-Buchoux M., Beninger PG , Chadenas C, Sellier D. 2017. European seabirds show stable contemporary biogeography. Waterbirds 40(4): 309-321   pdf

96 - Beninger PG. 2017. Caveat observator: the many faces of pre-spawning atresia in marine bivalve reproductive cycles. Mar Biol 164:163  pdf

95 - Dagorn FA, Couzinet-Mossion A, Kendel M, Beninger PG , Rabesaotra V, Barnathan G, Wielgosz-Collin G. 2016. Exploitable lipids and fatty acids in the invasive oyster Crassostrea gigas on the French Atlantic Coast. Mar. Drugs 14 (16): 104  pdf

94 - Beninger PG, Valdizan A, Le Pennec G. 2016. The seminal receptacle and implications for reproductive processes in the invasive gastropod Crepidula fornicata. Zoology 119:4-10  pdf

93 - Boldina I, Beninger PG. 2016. Strengthening statistical usage in marine ecology: Linear regression. J. Exp. Mar. Biol. Ecol 474:81-91  pdf

92 - Beninger PG, Boldina I. 2014. Fine-scale spatial distribution of the temperate infaunal bivalve Tapes (=Ruditapes) philippinarum (Adams and Reeve) on fished and unfished intertidal mudflats. J. Exp. Mar. Biol. Ecol 457:128-134 pdf

91 - Boldina I, Beninger PG. 2014. Fine-scale spatial distribution of the common lugworm Arenicola marina, and effects of intertidal clam fishing. Estuar. Coast. Shelf Sci 143: 32-40  pdf

90 - Boldina I, Beninger PG, Le Coz M. 2014. Effect of long-term mechanical perturbation on intertidal soft-bottom meiofunal community spatial structure.  J Sea Res 85: 85-91  pdf

89 - Boldina I, Beninger PG. 2013.  Fine-scale spatial structure of the exploited infaunal bivalve Cerastoderma edule on the French Atlantic coast.  J Sea Res 76: 193 – 200   pdf

88 – Beninger PG, Boldina I, Katsanevakis S. 2012.  Strengthening statistical usage in marine ecology.  J. Exp. Mar. Biol. Ecol 426-427: 97-108   pdf

87 – Valdizan A, Beninger PG, Decottignies P, Chantrel M, Cognie B.  2011. Evidence that rising coastal seawater temperatures increase reproductive output of the invasive gastropod Crepidula fornicata.  Mar. Ecol. Prog. Ser. 438: 153-165   pdf

86 - Beninger PG, Decottignies P, Elner RW.  2011.  Downward trophic shift during breeding migration in the shorebird  Calidris mauri (Western sandpiper).  Mar. Ecol. Prog. Ser. 428: 259-269   pdf

85 - Dutertre M,  Beninger PG,  Barillé L, Papin M, Haure J.  2010.  Rising water temperatures, reproduction and recruitment of an invasive oyster, Crassostrea gigas, on the French Atlantic coast.  Mar. Env. Res. 69: 1-9   pdf

84 - Beninger PG, Valdizan A, Decottignies P, Cognie B.  2010.  Field reproductive dynamics of the invasive slipper limpet, Crepidula fornicata. J. Exp. Mar. Biol. Ecol. 390:179–187   pdf

83 - Beninger PG.  2009.  Introduction to ‘Directions in Bivalve feeding’.  Aquat. Biol. 6: 175-180   pdf

82 - Yahel G, Marie D, Beninger PG, Eckstein S, Genin A.  2009.  In situ evidence for pre-capture qualitative selection in the tropical bivalve Lithophaga simplex.  Aquat. Biol. 6: 236 - 246   pdf

81 - Dutertre M, Barillé L,  Beninger PG,  Rosa P, Gruet Y. 2009. Variations in the pallial organ sizes of the invasive oyster, Crassostrea gigas, along an extreme turbidity gradient.  Estuar. Coast. Shelf Sci. 85: 431-436  pdf

80 - Valdizan A, Beninger PG, Cognie B, Decottignies P.  2009.  External fertilization and excapsular development in Crepidula fornicata: evaluating the risk of invasion control by dredging, crushing, and on-site rejection.  Aquat. Living Res. 22: 1-8   pdf

79 - Dutertre M, Beninger PG, Barillé L, Papin M, Rosa P, Barillé AL, Haure J. 2009.   Temperature and seston quantity and quality effects on field reproduction of farmed oysters, Crassostrea gigas, in Bourgneuf Bay, France.  Aquat. Liv. Res. 22: 319-329   pdf

78 - Cannuel R., Beninger PG,McCombie H, Boudry P. 2009.  Gill development and its functional and evolutionary implications in the blue mussel Mytilus edulis (Bivalvia: Mytilidae).   Biol. Bull.  217: 173-188   pdf

77 - Beninger PG, Valdizan A, Cognie B, Guiheneuf F, Decottignies P.  2008. Wanted: alive and not dead – functioning diatom status is a quality cue for the suspension-feeder Crassostrea gigas.  J. Plankton Res. 30: 689 - 697   pdf

76 - Beninger PG, Cognie B, Valdizan A, Decottignies P.  2008.  Impact of seston characteristics on qualitative particle selection sites and efficiencies in the pseudolamellibranch bivalve Crassostrea gigas.  J. Exp. Mar. Biol. Ecol. 360: 9-14   pdf

75 - Kuwae T, Beninger PG, Decottignies P, Mathot KJ, Lund DR, Elner RW. 2008. Biofilm grazing in a higher vertebrate: the Western sandpiper.  Ecology 89: 599-606   pdf

74 - Beninger PG, Decottignies P. 2008. Worth a second look: gill structure in Hemipecten forbesianus Adams & Reeve, and taxonomic implications for the Pectinidae.  J Mollusc Stud 74: 137-142   pdf

73 – Cannuel R, Beninger PG.  2007.  Acquisition of particle processing capability in juvenile oyster Crassostrea gigas: ontogeny of gill mucocytes.  Mar. Biol. 151: 897-905   pdf

72 - Decottignies P, Beninger PG, Rincé Y, Riera P.  2007. Trophic interactions between two introduced suspension-feeders, Crepidula fornicata and Crassostrea gigas, are influenced by seasonal effects and qualitative selection capacity. J. Exp. Mar. Biol. Ecol., 342 : 231-241   pdf

71 – Decottignies P, Beninger PG, Rincé Y, Robins RJ, Riera P. 2007. Exploitation of natural food sources by two sympatric, invasive suspension-feeders, Crassostrea gigas and Crepidula fornicata in Bourgneuf Bay, France.   Mar. Ecol. Prog. Ser. 334: 179-192   pdf

70 - Beninger PG, Decottignies P, Guiheneuf F, Barillé L, Rincé Y. 2007. Comparison of particle processing by two introduced  suspension-feeders: selection in Crepidula fornicata and Crassostrea gigas.  Mar. Ecol. Prog. Ser. 334: 165-177 pdf 

69 - Barillé L, Cognie B, Beninger PG, Decottignies P, Rincé Y. 2006. Feeding responses of the gastropod Crepidula fornicata in turbid environments.  Mar. Ecol. Prog. Ser. 322: 169-178   pdf

68 – Beninger PG, Cannuel R. 2006.  Acquisition of particle-processing capability in the postlarval oyster Crassostrea gigas: ontogeny of the mantle pseudofeces rejection tracts.  Mar. Ecol. Prog. Ser. 325: 153 - 163  pdf

67 – Cannuel R, Beninger PG. 2006. Gill development, functional and evolutionary implications in the Pacific oyster Crassostrea gigas (Bivalvia: Ostreidae).  Mar. Biol. 149: 547 – 563  pdf

66 - Beninger PG, Le Pennec M.  2006.  Structure and function in scallops.  In: S.E. Shumway and J. Parsons (eds.), Scallops: Biology, ecology and aquaculture (2nd edition).  Elsevier science publishers, Amsterdam,  pp 123 - 227

65 - Dufour SC, Steiner G, Beninger PG.  2006.  Phylogenetic analysis of the peri-hydrothermal vent bivalve Bathypecten vulcani based on 18S rRNA.  Malacologia 48: 35-42   pdf

64 – Dubois S, Barillé L, Cognie B, Beninger PG. 2005. Particle capture and processing mechanisms in Sabellaria alveolata (Polychaeta: Sabellariidae).  Mar. Ecol. Prog. Ser. 301: 159 - 171   pdf

63 – Beninger PG, Cannuel R, Jaunet S.  2005.  Particle processing on the gill plicae of the oyster Crassostrea gigas: fine-scale mucocyte distribution and functional correlates.  Mar. Ecol. Prog. Ser. 295: 191 - 199   pdf

62 – Cannuel R, Beninger PG.  2005.  Is oyster broodstock feeding always necessary?  A study using oocyte quality predictors and validators in Crassostrea gigas.  Aquatic Living Res. 18: 35-43   pdf

61 - Beninger PG, Decottignies P. 2005. What makes diatoms attractive to suspensivores? The organic casing and associated organic molecules of Coscinodiscus perforatus are quality cues for the  bivalve Pecten maximus.   J. Plankton Biol. 27: 11-17   pdf

60 - Elner RW, Beninger PG, Jackson DL, Potter T. 2005. Evidence for a new feeding mode in Western Sandpiper (Calidris mauri ) and Dunlin (Calidris alpina) based on bill and tongue morphology and ultrastructure.  Mar. Biol. 146: 1223-1234   pdf

59 - Beninger PG, Decottignies P, Rincé Y.  2004.  Localization of qualitative selection sites in the heterorhabdic filibranch Pecten maximus L. (Bivalvia: Pectinidae).  Mar. Ecol. Prog. Ser. 275: 163 - 173   pdf

58 - Beninger PG,  Le Pennec G, Le Pennec, M. 2003.  Demonstration of nutrient pathway from the digestive system to oocytes in the gonad intestinal loop of the scallop Pecten maximus L.  Biol. Bull. 205: 83 - 92   pdf

57 - Cognie B, Barillé L, Massé G, Beninger PG.  2003.  Selection and processing of large suspended algae in the oyster Crassostrea gigas. Mar. Ecol. Prog. Ser.  250: 145 – 152   pdf

56 - Le Pennec, G.,  Beninger PG, Le Pennec M,  Donval A.  2003.  Aspects of the feeding biology of the pectinacean Bathypecten vulcani, a perihydrothermal vent bivalve.  J. Mar. Biol.Assoc. UK 83: 479 - 482   pdf

55 - Veniot A, Bricelj M, Beninger PG.  2003. Ontogenetic changes in gill morphology and potential significance for food acquisition in the scallop Placopecten magellanicus. Mar. Biol. 142: 123 - 131   pdf

54 - Beninger PG, Dufour SC, Decottignies P, Le Pennec M. 2003.  Particle processing mechanisms in the archaic, peri-hydrothermal vent bivalve Bathypecten vulcani, inferred from cilia and mucocyte distributions on the gill.  Mar. Ecol Prog. Ser. 246: 183 – 195   pdf

53 – Le Pennec G, Le Pennec M, Beninger PG, Dufour SC.  2002.  Spermatogenesis in the archaic hydrothermal vent bivalve, Bathypecten vulcani, and comparison of spermatozoon ultrastructure with littoral pectinids.  Inv. Reprod. Devel. 41: 13 - 19   pdf

52  – Le Pennec G, Le Pennec M, Beninger PG.  2001. Seasonal digestive gland dynamics of the scallop Pecten maximus L in the Bay of Brest (France).  J. Mar. Biol. Ass. UK 81: 663 – 671   pdf

51 - Beninger PG, Cannuel R, Blin J-L Pien S, Richard O.  2001.  Reproductive characteristics of the archaeogastropod Megathura crenulata.  J. Shellfish Res. 20: 301 - 307   pdf

50 - Dufour SC, Beninger PG.  2001.  A functional interpretation of cilia and mucocyte distributions on the abfrontal surface of bivalve gills.  Mar. Biol., 138: 295 - 309   pdf

49 - Beninger PG, Dufour SC.  2000.  Evolutionary trajectories of a redundant feature: lessons from bivalve gill  abfrontal cilia and mucocyte distributions.  In: Harper, E.M., Taylor, J.D., & Crame, J.A. (eds) The Evolutionary Biology of the Bivalvia.  Geological Society of London Special Publication:  273 – 278

48 - Beninger PG. 2000. Limits and constraints: a comment on premises and methods in recent studies concerning particle capture mechanisms in bivalves. Limnol. Oceanogr. 45: 1196 - 1199   pdf

47 - Le Pennec M,  Beninger PG. 2000.  Reproductive characteristics and strategies of reducing-system bivalves.  Comp. Biochem. Physiol. 126A: 1 – 16   pdf

46 - Silverman H, Lynne JW, Beninger PG, Dietz TH.  1999.  The role of latero-frontal cirri in particle capture by the gills of Mytilus edulisBiol. Bull.197: 368 – 376   pdf

45 - Beninger PG, Veniot A.  1999.  The oyster proves the rule: mechanisms of pseudofeces transport and rejection in Crassostrea virginica and C. gigas.  Mar. Ecol Prog. Ser. 190: 179 - 188   pdf

44 - Beninger PG, Veniot A, Poussart Y. 1999.  Principles of pseudofeces rejection on the bivalve mantle:  integration in particle processing.  Mar. Ecol Prog. Ser. 178: 259 - 269   pdf

43 - Beninger PG, Larocque R. 1998. Gonopod tegumental glands: a new accessory sex gland in the Brachyura.  Mar. Biol. 132: 435-444   pdf

42 - Beninger PG, Saint-jean S. 1997. Particle processing on the labial palps of Mytilus edulis and Placopecten magellanicus (Mollusca, bivalvia). Mar. Ecol. Progr. Ser. 147: 117-127   pdf

41 - Le Pennec M, Beninger PG. 1997. Ultrastructural characteristics of spermatogenesis in three species of deep-sea hydrothermal vent mytilids.  Can. J. Zool. 75: 308-316   pdf

40 - Beninger PG, Le Pennec M. 1997. Reproductive characteristics of a primitive bivalve from a deep-sea reducing environment: giant gametes and their significance in Acharax alinae (Cryptodonta: Solemyidae).  Mar. Ecol. Progr. Ser. 157: 195-206   pdf

39 - Beninger PG, Lynn JW, Dietz TH, Silverman H. 1997. Mucociliary transport in living tissue: the two-layer model confirmed in the mussel Mytilus edulis L.  Biol. Bull. 193: 4-7   pdf

38 - Beninger PG, Saint-Jean S. 1997.  The role of mucus in particle processing by suspension-feeding marine bivalves: unifying principles.  Mar. Biol. 129: 389-397   pdf

37 - Beninger PG, Dufours S, Bourque J. 1997.  Particle processing mechanisms of the eulamellibranchs Spisula solidissima and Mya arenaria.  Mar Ecol. Prog. Ser. 150: 157-169   pdf

36 - Beninger PG, Dufours S. 1996. Mucocyte distribution and relationship to particle transport on the pseudolamellibranch gill of Crassostrea virginica (Bivalvia: Ostreidae).  Mar. Ecol. Prog. Ser. 137: 133-138   pdf

35 – Lanteigne C, Beninger PG, Gionet C. 1996.  Ontogeny of female primary sexual characters in the majid crab Chionocetes opilio and Hyas coarctus.  J. Crust. Biol. 16: 501-514   pdf

34 - Beninger PG, Ferguson A, Lanteigne C. 1995.  The gonopod tegumental glands of snow crab Chionoecetes opilio are accessory reproductive glands.  J. Shellfish Res. 14: 365-370   pdf

33 - Elner RW, Beninger PG. 1995. Multiple reproductive strategies in snow crab Chionoecetes opilio: physiological pathways and behavioral plasticity.  J. Exp. Mar. Biol. Ecol. 193: 93-112   pdf

32 - Beninger PG, Saint-Jean S, Poussart Y. 1995.  Labial palps of the blue mussel, Mytilus edulis L. (Bivalvia, Mytilidae).  Mar. Biol. 123: 293-303   pdf

31 – Le Pennec M., Beninger PG, Herry A. 1995.  Feeding and digestive adaptations of bivalve molluscs to sulphide-rich habitats.  Comp. Biochem. Physiol. 111A: 183-189   pdf

30 - Beninger PG, Potter TM, Saint-Jean S. 1995. Paddle cilia fixation artefacts in pallial organs of adult Mytilus edulis and Placopecten magellanicus (Mollusca, Bivalvia).  Can. J. Zool. 73: 610-614   pdf

29 - Beninger PG, Donval A, Le Pennec M. 1995.  The osphradium in Placopecten magellanicus and Pecten maximus (Bivalvia, Pectinidae): histology, ultrastructure and implications for spawning synchronisation. Mar. Biol. 123: 121-129   pdf

28 – Le Pennec M, Herry A, Johnson M, Beninger PG. 1995.  Nutrition-gametogenesis relationship in the endosymbiont host bivalve Loripes lucinalis (Lucinidae) from reducing coastal habitats. - In :  Eleftheriou, A., Ansell, A.D. & Smith, C.J. (eds) Biology and ecology of shallow coastal waters. Proc. 28th European Mar. Biol. Symp., Olsen and Olsen Co: 139-142

27 - Beninger PG, Dwiono S, Le Pennec M. 1994. Early development of the gill and implications for feeding in Pecten maximus (Bivalvia : Pectinidae). Mar. Biol. 119: 405–412   pdf

26 - Beninger PG, St-Jean S, Poussart Y, Ward E. 1993.  Gill function and mucocyte distribution in Placopecten magellanicus and Mytilus edulis (Mollusca : Bivalvia) :  the role of mucus in particle transport.  Mar. Ecol. Prog. Ser. 98: 275–282   pdf

25 - Beninger PG, Le Pennec M.  1993.  A histochemical study of the bucco-oesophageal glands of Mytilus edulis: the importance of mucus in ingestion.  J. Mar. Biol. Assoc. U.K. 73:237-240   pdf

24 - Ward JE, MacDonald BA, Thompson RJ, Beninger PG. 1993. Mechanisms of suspension-feeding in Bivalves : resolution of current controversies using endoscopy.  Limnol. Oceanogr. 30 : 265-272   pdf

23 - Beninger PG, Lanteigne C, Elner RW. 1993. Reproductive processes revealed by spermatophore dehiscence experiments and by histology, ultrastructure, and histochemistry of the female reproductive system in the snow crab, Chionoecetes opilio (O. Fabricius).  J. Crust. Biol. 13: 1-16   pdf

22 - Le Pennec M, Martinez JC, Donval A, Herry A, Beninger PG. 1992. Enzymologie du tractus digestif de la Modiole hydrothermale Bathymodiolus thermophilus.  Can. J. Zool. 2298-2302   pdf

21 - Beninger PG, Ward J E, MacDonald BA, Thompson RJ.  1992. Gill function and particle transport in Placopecten magellanicus (Gmelin) (Mollusca : Bivalvia) as revealed using video endoscopy.  Mar. Biol. 114: 281-288   pdf

20 - Elner RW, Beninger PG.  1992. The reproductive biology of snow crab, Chionoecetes opilio : a synthesis of recent contributions.  Amer. Zool. 32: 524-533   pdf

19 - Le Pennec M, Dorange G, Beninger PG, Donval A,  Widowati I.  1991. Les relations trophiques anse intestinale-gonade chez Pecten maximus (Mollusque, Bivalve).  Haliotis 21: 57-69  

18 - Ward JE, Beninger PG, MacDonald BA, Thompson RJ.  1991.  Direct observations of feeding structures and mechanisms in Bivalve molluscs using endoscopic examination and video image analysis. Mar. Biol. 111: 287–291   pdf

17 - Beninger PG, Le Pennec M, Donval A. 1991.  Mode of particle ingestion in five species of Bivalve molluscs. Mar. Biol. 108: 255-261   pdf

16 - Le Pennec M, Beninger PG, Dorange G, Paulet YM. 1991. Trophic sources and pathways to the developing gametes of Pecten maximus (Bivalvia : Pectinidae). J. Mar. Biol. Ass. U.K. 71: 451-463   pdf

15 - Beninger PG.  1991.  Structures and mechanisms of feeding in scallops :  paradigms and paradoxes. In, S.E. Shumway (ed.) An International Compendium of Scallop Biology and Culture. World Aquaculture Workshops 1: 331-340

14 - Beninger PG, Elner RW, Poussart Y.  1991.  The gonopods of the majid crab Chionoecetes opilio (O. Fabricius). J. Crust. Biol. 11: 217-228 pdf

13 - Beninger PG, Auffret M, Le Pennec, M. 1990.  The peribuccal organs of Placopecten magellanicus and Chlamys varia (Mollusca :  Bivalvia) : structure, ultrastructure and implications for feeding. I. The labial palps.  Mar. Biol. 107: 215-223   pdf

12 - Beninger PG, Le Pennec M, Auffret M. 1990.  The peribuccal organs of Placopecten magellanicus and Chlamys varia (Mollusca : Bivalvia) : structure, ultrastructure, and implications for feeding. II. The lips.  Mar. Biol. 107: 225-233   pdf

11 - Beninger PG, Elner RW, Foyle TP, OdensePH. 1988.  Functional anatomy of the male reproductive system and the female spermatheca in the snow crab, Chionoecetes opilio (O. Fabricius) (Decapoda : Majidae) and a hypothesis for fertilization.  J. Crust. Biol. 8: 322-332  pdf

10 - Beninger PG, Le Pennec M, Salaün M. 1988.  New observations of the gills of Placopecten magellanicus Gmelin (Mollusca : Bivalvia), and implications for nutrition.  I. General anatomy and surface micro-anatomy.  Mar. Biol. 98: 61-70   pdf

9 - Le Pennec M, Beninger PG, Herry A. 1988.  New observations of the gills of Placopecten magellanicus Gmelin (Mollusca :  Bivalvia), and implications for nutrition.  II. Internal anatomy and micro-anatomy.  Mar. Biol. 98: 229-237   pdf

8 - Beninger PG. 1987.  A qualitative and quantitative study of the reproductive  cycle of the giant scallop Placopecten magellanicus in the Bay of Fundy.  (New Brunswick, Canada).  Can. J. Zool. 65: 495-498  pdf

7 - Beninger PG, Chiasson L, Elner RW. 1986.  The utility of artificial collectors as a technique to study benthic settlement and early juvenile growth of the rock crab, Cancer irroratus.  Fish. Res., 4: 317-329   pdf

6 - Lucas A. Chebab-Chalabi L, and Beninger PG. 1986.  Variation of relative organic matter in Mytilus edulis (L.) larvae and postlarvae. J. Exp. Mar. Biol. Ecol. 95: 99-103   pdf

5 - Beninger PG, Stephan G. 1985.  Seasonal variations in the fatty acids of the triacylglycerols and phospholipids of two populations of adult clam (Tapes decussatus L. and Tapes philippinarum) reared in a common habitat.  Comp. Biochem. Physiol. 81 B: 591-601   pdf

4 - Beninger PG. 1985.  Long-term variations in cation content of two populations of adult marine clam (Tapes decussatus L. and T. philippinarum) reared in a common habitat.  Comp. Biochem. Physiol. 82 A: 945-949   pdf

3 - Lucas A, Beninger PG. 1985.  The use of physiological condition indices in marine Bivalve aquaculture.  Aquaculture 44: 187-200   pdf

2 - Beninger PG. 1984.  Seasonal variations of the major lipid classes in relation to the reproductive activity of two species of clams raised in a common habitat :  Tapes decussatus L. (Jeffreys, 1863) and Tapes philippinarum (Adams et Reeve, 1850).  J. Exp. Mar. Biol. Ecol. 79: 79-90   pdf

1 - Beninger PG, Lucas A.  1984.  Seasonal variations in condition, reproductive activity, and gross biochemical composition of two species of adult clam reared in a common habitat :  Tapes decussatus L. (Jeffreys) and Tapes philippinarum (Adams and Reeve).  J. Exp. Mar. Biol.Ecol. 79: 19-38   pdf

2. Books and Book Chapters

10 - Beninger PG (editor). 2018. Mudflat Ecology. Springer Nature Publishers, Switzerland, i-xiv+ 429pp. 

9 - Beninger PG, Patterson DM. 2018. Chapter 1. Introduction: Mudflat Basics. In Beninger, PG (ed), Mudflat Ecology. Springer Nature Switzerland: 1-10.

8 - Moens T, Beninger PG. 2018. Chapter 5. Meiofauna: an inconspicuous but important player in mudflat ecology. In Beninger, PG (ed), Mudflat Ecology. Springer Nature Switzerland: 91-112.

7 - Beninger PG, Cuadrado D, Van de Koppel J. 2018. Chapter 8. Sedimentary and biological patterns on mudflats. In Beninger, PG (ed), Mudflat Ecology. Springer Nature Switzerland: 185-212.

6 - Beninger PG. 2018. Chapter 13. Mudflat Fishing. In Beninger, PG (ed), Mudflat Ecology. Springer Nature Switzerland: 339-365.

5 - Beninger PG, Shumway SE. 2018. Chapter 14. Mudflat Aquaculture. 2018. In Beninger, PG (ed), Mudflat Ecology. Springer Nature Switzerland: 365-388. pdf

4 - Beninger PG, Boldina I. 2018. Chapter 15. Quantitative considerations in mudflat ecology. In Beninger, PG (ed), Mudflat Ecology. Springer Nature Switzerland: 389-418.

3 - Beninger PG, Le Pennec M. 2016. Chapter 3. Scallop structure and function. In Shumway SE, Parsons GJ (eds), Scallops: Biology, Ecology, Aquaculture, and Fisheries, 3rd edition, Elsevier Science: 85-159. pdf

2 - Beninger PG and M. Le Pennec, 2006.  Structure and function in scallops.  In S. Shumway and GJ Parsons (editors), Scallops : Their Biology, Ecology and Aquaculture.  2nd edition.  Elsevier Science Publishers, Amsterdam: 123-227. 

1 - Beninger PG and M. Le Pennec, 1991.  Functional anatomy of scallops.  In S. Shumway (editor), Scallops : Their Biology, Ecology and Aquaculture.  Elsevier Science Publishers, Amsterdam: 133-223. 

3. Comments and letters in refereed journals

11 - Beninger PG . 2023. Book Review. New horizons in meiobenthos research. By Olav Giere, Michaela Schratzberger (Eds.).
Invertebrate Biology. 142(2)  pdf

10 - Beninger PG ,Chérel D, Le Pennec G, Shumway SE. 2022. Bivalve oocyte inviability revealed using a simple staining technique.
ECOSHERE. 13(6)  pdf

9 - Eveillard-Buchoix M, Beninger PG. 2022. Stepping stones to reproductive success: The role of geomorphology in seabird nest site choice.
Bulletin of the Ecological Society of America. 103(1)  pdf

8 - Eveillard-Buchoix M, Beninger PG. 2022. Ecology 2022. Cover Image. Bulletin of the Ecological Society of America. 103(1) pdf

7 - Beninger PG, Elner R.W. 2020. TONGUE IN PEEP: NEW PERSPECTIVES FOR SHOREBIRD ECOLOGY. Bulletin of the Ecological Society of America. 101(3) pdf

6 - Beninger PG, Chérel D. 2019. What cloaked bivalve oocytes teach us about reproductive biology, evolution, and scientific awareness. Bulletin of the Ecological Society of America. 101(1) pdf

5 - Bayne B, Anglès d'Auriac M, Backeljau T, Beninger PG, Boudry P, Carnegie R , Davis J, Guo X , Hedgecock D, Krause M, Langdon C, Lapègue S, Manahan D , Mann R, Powell E, Shumway S. 2019. A scientific name for Pacific oysters. Aquaculture 499: 373. pdf

4 - Bayne BL, Ahrens M, Allen SK, Anglès D'Auriac M, Backeljau T, Beninger PG, Bohn R, Boudry P, Davis J, Green T, Guo X, Hedgecock D, Ibarra A, Kingsley-Smith P, Krause M, Langdon C, Lapègue S, Li C, Manahan D, Mann R, Perez-Paralle L, Powell EN, Rawson PD, Speiser D, Sanchez J-L, Shumway S, Wang H. 2017. The proposed dropping of the genus Crassostrea for all Pacific cupped oysters and its replacement by a new genus Magallana: a dissenting view. J.Shellfish Res. 36: 545-547. pdf

3 - Beninger PG, Beall J, Shumway SE. 2016. Debasing the currency of science: the growing menace of predatory open access journals. J. Shellfish Res. 35: 1–5. pdf

2 - Ward JE, MacDonald BA, Thompson RJ, Beninger PG. 1993. The role of mucus in bivalve feeding - reply to comment by C.B. Jørgensen. Limnol. Oceanogr. 38: 467.   pdf

1 - Elner RW, Beninger PG. 1989. Functional maturity in small snow crabs (Chionoecetes opilio): sizing up the evidence. Can. J. Fish. Aquat. Sci., 46: 2037‑2039.

4. Technical report

1 - Elner RW, Beninger PG., Linkletter LE, Lanteigne S. 1985. Guide to indicator fragments of principal prey taxa in the stomac s of two common atlantic crab species: Cancer borealis Stimpson, 1859 and Cancer irroratus Say, 1817. Canadian Technical Report of Fisheries and Aquatic Sciences. 1403.  pdf